Lastly, N is converted to NO by energy from lightning, which is negligible in current temperate ecosystems, or by fossil fuel combustion.

The sanctuary represents the easternmost temperate ecosystems and landscapes of Bhutan, and is part of the Eastern Himalayan subalpine conifer forests ecoregion.

Species diversity, nutrient availability, precipitation, and average temperatures increase as you move from the tundra to boreal forests and then to deciduous temperate ecosystems, which are found south of these Arctic biomes.

It is a frequent colonizer of temperate riparian ecosystems, roadsides and waste places.

According to UNESCO "The area covered by the World Heritage site is claimed to be the most biodiverse and least disturbed temperate ecosystems in the world".

The east side of the municipality is dominated by temperate ecosystems, especially in the higher altitudes.

Marking a transition zone between tropical, temperate and semidesert ecosystems, its diversity is incredible.

Ectomycorrhizal fungi are found throughout boreal, temperate and tropical ecosystems, primarily among the dominant woody-plant-producing families (3.1.

This is far longer than in temperate ecosystems.

Unesco said the region "may be the most biologically diverse temperate ecosystem in the world" and designated it a World Heritage Site in the summer of 2003.