His research interests are micro-RNA responses to radiation exposure, radiation-induced bystander effect, cell cycle perturbations and the enzymatic processing of clustered DNA damage.

The Radiation-Induced Bystander Effect (Bystander Effect) is the phenomenon in which unirradiated cells exhibit irradiated effects as a result of signals received from nearby irradiated cells.

The prominent theme of research undertaken using the RARAF microbeam is damage signal transduction, both within cells and between cells, which is of interest due in part to the discovery of the radiation-induced bystander effect.