Lardacein is a protein found in tissues affected with amyloid degeneration.

Oettingen is credited for providing the first comprehensive description of amyloid degeneration of the eye's conjunctiva.

Amyloid degeneration is a type of degeneration with the deposit of lardacein in the tissues.

During this time he completed his first scientific works, and he was awarded the Gold Medal for research on the "Histogenesis of experimentally induced amyloid degeneration of the liver in animals" published in the journal Russian Archives of pathology, clinical medicine and bacteriology.